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{{Taxobox| color = pink| name = Dolphin| image = Bottlenose_Dolphin_KSC04pd0178.jpg| image_width = 250px| image_caption =
Bottlenose Dolphin breaching in the bow wave of a boat] - Recent| regnum =
Animalia]| classis =
Mammalia]| subordo =
Odontoceti, 1821| subdivision_ranks = [Genus| subdivision =See article below.-->
Dolphins are aquatic mammals that are closely related to whales and
porpoises. There are almost forty species of dolphin in seventeen genus. They vary in size from 1.2
metres (4
foot (unit of length)) and 40 kilograms (88
pound (unit of weight)) (Maui's Dolphin), up to 9.5 m (30 ft) and ten tonnes (the Orca or Killer Whale). They are found worldwide, mostly in the shallower seas of the
continental shelves, and are carnivores, mostly eating fish and squid. The
Family (biology) Oceanic dolphin is the largest in the Cetacea, and relatively recent: dolphins evolved about ten million years ago, during the Miocene. Dolphins are considered to be amongst the most intelligent of animals and their often friendly appearance and seemingly playful attitude have made them popular in human culture.
Origin of the name
The name is originally from Ancient Greek (
delphís; "dolphin"), which was related to the Greek (
delphys; "womb"). The animal's name can therefore be interpreted as meaning "a 'fish' with a womb".The American Heritage® Dictionary of the English Language, Fourth Edition, online entry at Dictionary.com, retrieved
December 17 2006. The name was transmitted via the
Latin delphinus, Middle Latin
dolfinus and the
Old French daulphin, which reintrodued the
ph into the word.
The word is used in a few different ways. It can mean:
- Any member of the family Delphinidae (oceanic dolphins),
- Any member of the families Delphinidae and Platanistoidea (oceanic and river dolphins),
- Any member of the suborder Odontoceti (toothed whales; these include the above families and some others),
- Used casually as a synonym for Bottlenose Dolphin, the most common and familiar species of dolphin.
In this article, the second definition is used. Porpoises (suborder Odontoceti, family
Phocoenidae) are thus not dolphins in this sense. Orcas and some closely related species belong to the Delphinidae family and therefore qualify as dolphins, even though they are called whales in common language. A group of dolphins can be called a "school" or a "pod".
Taxonomy
- Suborder Odontoceti, toothed whales
- Family Delphinidae, oceanic dolphins
- Genus Delphinus
- Genus Tursiops
- Bottlenose Dolphin, Tursiops truncatus
- Indo-Pacific Bottlenose Dolphin, Tursiops aduncus
- Genus Lissodelphis
- Northern Rightwhale Dolphin, Lissodelphis borealis
- Southern Rightwhale Dolphin, Lissiodelphis peronii
- Genus Sotalia
- Genus Sousa
- Genus Stenella
- Atlantic Spotted Dolphin, Stenella frontalis
- Clymene Dolphin, Stenella clymene
- Pantropical Spotted Dolphin, Stenella attenuata
- Spinner Dolphin, Stenella longirostris
- Striped Dolphin, Stenella coeruleoalba
- Genus Steno
- Rough-Toothed Dolphin, Steno bredanensis
- Genus Cephalorynchus
- Genus Grampus
- Genus Lagenodelphis
- Fraser's Dolphin, Lagenodelphis hosei
- Genus Lagenorhyncus
- Atlantic White-Sided Dolphin, Lagenorhynchus acutus
- Dusky Dolphin, Lagenorhynchus obscurus
- Hourglass Dolphin, Lagenorhynchus cruciger
- Pacific White-Sided Dolphin, Lagenorhynchus obliquidens
- Peale's Dolphin, Lagenorhynchus australis
- White-Beaked Dolphin, Lagenorhynchus albirostris
- Genus Orcaella
- Australian Snubfin Dolphin, Orcaella heinsohni
- Irrawaddy Dolphin, Orcaella brevirostris
- Genus Peponocephala
- Melon-headed Whale, Peponocephala electra
- Genus Orcinus
- Killer Whale, Orcinus orca
- Genus Feresa
- Genus Pseudorca
- Genus Globicephala
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhynchus
- Family Platanistoidea, river dolphins
- Genus Inia
- Boto (Amazon River Dolphin), Inia geoffrensis
- Genus Lipotes
- Chinese River Dolphin (Baiji), Lipotes vexillifer (considered functionally extinct)
- Genus Platanista
- Genus Pontoporia
- La Plata Dolphin (Franciscana), Pontoporia blainvillei
Six species in the family Delphinidae are commonly called "whales" but are strictly speaking dolphins. They are sometimes called "blackfish".
- Melon-headed Whale, Peponocephala electra
- Orca, Orcinus orca
- Pygmy Killer Whale, Feresa attenuata
- False Killer Whale, Psudorca crassidens
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhynchus
Hybrid dolphins
In 1933, three abnormal dolphins were beached off the Ireland coast; these appeared to be
hybrids between Risso's Dolphin and the Bottlenose Dolphin.Dolphin Safari (2006) sightings log, page retrieved December 17 2006. This mating has since been repeated in captivity and a hybrid calf was born. In captivity, a Bottlenose Dolphin and a Rough-toothed Dolphin produced hybrid offspring.Texas Tech University (1997), Mammals of Texas - Rough-toothed Dolphin, article retrieved December 8
2006. A Common-Bottlenose hybrid lives at SeaWorld California.Robin's Island Dolphins at SeaWorld California, page retrieved December 17
2006. Various other dolphin hybrids have also been reported in the wild, such as a Bottlenose-Atlantic Spotted hybrid.Denise L. Herzing, Kelly Moewe and Barbara J. Brunnick (2003), Interspecies interactions between Atlantic spotted dolphins, Stenella frontalis and bottlenose dolphins, Tursiops truncatus, on Great Bahama Bank, Bahamas, article retrieved on
December 17 2006. The best known hybrid however is the Wolphin, a False Killer Whale-Bottlenose Dolphin hybrid. The Wolphin is a fertile hybrid, and two such Wolphins currently live at the Sea Life Park in
Hawaii, the first having been born in 1985 from a male False Killer Whale and a female Bottlenose. Wolphins have also been observed in the wild.Louis Herman, interviewed for Associated Press, article by Jeanette J. Lee (2005), Livescience.com - Whale-Dolphin Hybrid Has Baby Wholphin, article retrieved April 26, 2007.
Evolution and anatomy
Evolution
Dolphins, along with whales and porpoises, are thought to be descendants of terrestrial mammals, most likely of the
Artiodactyl order (biology). The ancestors of the modern day dolphins entered the water roughly fifty million years ago, in the Eocene epoch.
Modern dolphin
skeletons have two small, rod-shaped pelvic bones thought to be
vestigial hind legs. In
October 2006 an unusual Bottlenose Dolphin was captured in
Japan; it had small fins on each side of its genital slit which scientists believe to be a more pronounced development of these vestigial hind legs.Associated Press /
FOX news (2006), Japanese Researchers Find Dolphin With 'Remains of Legs', article retrieved
November 6 2006.
Anatomy
Dolphins have a streamlined
fusiform body, adapted for fast swimming. The basic colouration patterns are shades of grey with a light underside and a distinct dark cape on the back. It is often combined with lines and patches of different hue and contrast.
The head contains the
melon (whale), a round organ used for animal echolocation. In many species, the jaws are elongated, forming a distinct beak; for some species like the Bottlenose, there is a curved mouth which looks like a fixed smile. Teeth can be very numerous (up to two hundred and fifty) in several species. The dolphin brain is large and has a highly structured cortex, which often is referred to in discussions about their advanced intelligence.
Unlike most mammals, dolphins do not have hair, but they are born with a few hairs around the tip of their
rostrum (anatomy) which they lose after some time, in some cases even before they are born. The only exception to this is the Boto river dolphin, which does have some small hairs on the rostrum.
Their reproductive organs are located on the underside of the body. Males have two slits, one concealing the
penis and one further behind for the
anus. The female has one genital slit, housing the vagina and the anus. A mammary slit is positioned on either side of the female's genital slit.
Senses
Most dolphins have acute eyesight, both in and out of the water, and their sense of Hearing (sense) is superior to that of humans. Though they have a small ear opening on each side of their head, it is believed that hearing underwater is also if not exclusively done with the lower jaw which conducts the sound vibrations to the middle ear via a fat-filled cavity in the lower jaw bone. Hearing is also used for Animal_echolocation, which seems to be an ability all dolphins have. Their teeth are arranged in a way that works as an array or antenna to receive the incoming sound and make it easier for them to pinpoint the exact location of an object.Goodson, A.D. and M. Klinowska. "A Proposed Echolocation Receptor for the Bottlenose Dolphin (
Tursiops truncatus): Modelling the Receive Directivity from Tooth and Lower Jaw Geometry", in Thomas and Kastelein, eds,
NATO ASI Series A: Sensory Abilities of Cetaceans, vi.196:255-267 (Plenum NY, 1990) ISBN 0-30-643695-7 The dolphin's sense of touch is also well-developed. However, dolphins lack an olfactory nerve and lobes and thus are believed to have no sense of smell,SeaWorld, Bottlenose Dolphins - Senses, article retrieved
December 17 2006. but they can
taste and do show preferences for certain kinds of fish. Since dolphins spend most of their time below the surface normally, just tasting the water could act in a manner analogous to a sense of smell.
Though most dolphins do not have any hair, they do still have hair follicles and it is believed these might still perform some sensory function, though it is unclear what exactly this may be.Bjorn Mauck, Ulf Eysel and Guide Dehnhardt (2000), Selective heating of vibrissal follicles in seals (Phoca Vitulina) and dolphins (Sotalia Fluviatilis Guianensis), article retrieved
March 11, 2007. The small hairs on the rostrum of the Boto river dolphin are believed to function as a somatosensory system however, possibly to compensate for the Boto's poor eyesight.Laurie Stepanek (1998), Amazon River Dolphin (Inia geoffrensis), article retrieved
March 11, 2007.
Behaviour
Dolphins are often regarded as one of Earth's most intelligent animals, though it is hard to say just how intelligent dolphins are, as comparisons of species' relative intelligence are complicated by differences in sensory apparatus, response modes, and nature of cognition. Furthermore, the difficulty and expense of doing experimental work with large aquatics means that some tests which could yield meaningful results still have not been carried out, or have been carried out with inadequate sample size and methodology. Dolphin behaviour has been studied extensively by humans however, both in captivity and in the wild. See the cetacean intelligence article for more details.
Social behaviour
, Queensland, Australia.Dolphins are social, living in pods (also called "schools") of up to a dozen individuals. In places with a high abundance of food, pods can join temporarily, forming an aggregation called a
superpod; such groupings may exceed a thousand dolphins. The individuals communicate using a variety of clicks, whistles and other vocalizations. They also use ultrasonic sounds for echolocation. Membership in pods is not rigid; interchange is common. However, the cetaceans can establish strong bonds between each other. This leads to them staying with injured or ill individuals.
In May 2005, researchers in Australia discovered a cultural aspect of dolphin behaviour: Some dolphins (
Tursiops aduncus) teach their children to use tools. The dolphins break sea sponges off and cover their snouts with them thus protecting their snouts while foraging. This knowledge of how to use a tool is mostly transferred from mothers to daughters, unlike
simian primates, where the knowledge is generally passed on to both sexes. The technology to use sponges as mouth protection is not genetically inherited but a taught behaviour.Rowan Hooper for New Scientist (2005), Dolphins teach their children to use sponges, article retrieved
December 17 2006.
They are also occasionally willing to approach humans and playfully interact with them in the water. Dolphins have also been known to seemingly protect swimmers from sharks by swimming circles around them.CBC News (2004), Dolphins save swimmers from shark, article retrieved
March 11, 2007.
Dolphins are known to engage in acts of aggression towards each other. The older a male dolphin is, the more likely his body is covered with scars ranging in depth from teeth marks made by other dolphins. It is suggested that male dolphins engage in such acts of aggression for the same reasons as humans: disputes between companions or even competition for other females. Acts of aggression can become so intense that targeted dolphins are known to go into exile, leaving their communities as a result of losing a fight with other dolphins.
Male Bottlenose Dolphins have been known to engage in infanticide. Dolphins have also been known to kill
porpoises for reasons which are not fully understood, as porpoises generally do not share the same fish diet as dolphins and are therefore not competitors for food supplies.Dr. George Johnson (date unknown), Is Flipper A Senseless Killer?, article retrieved December 17 2006.
Reproduction and sexuality
Dolphin copulation happens belly to belly and though many species engage in lengthy
foreplay, the actual act is usually only brief, but may be repeated several times within a short timespan. The
gestation period varies per species; for the small Tucuxi dolphin, this period is around 11 to 12 months, while for the Orca the gestation period is around 17 months. They usually become sexually active at a young age already, even before reaching sexual maturity. The age at which sexual maturity is reached varies per species and gender.
Dolphins are known to have sex for reasons other than reproduction, sometimes also engaging in acts of a
homosexual nature.Herzing D.L., Rogers C.A., for the Wild Dolphin Project, Directionality of sexual aggression in mixed-species encounters between Atlantic Spotted dolphins and Bottlenose dolphins in the Bahamas (2005), article retrieved September 18, 2007. Various dolphin species have been known to engage in sexual behaviour with other dolphin species, this also having resulted in various hybrid dolphin species as mentioned earlier. Sexual encounters may be violent, with male dolphins sometimes showing aggressive behaviour towards both females and other male dolphins.Scott et. al, Aggression in bottlenose dolphins: evidence for sexual coercion, male-male competition, and female tolerance through analysis of tooth-rake marks and behaviour (2005), article retrieved September 18, 2007. Occasionally, dolphins will also show sexual behaviour towards other animals, including humans.Amy Samuels, Lars Bejder, Rochelle Constantine and Sonja Heinrich (2003), Marine Mammals: Fisheries, Tourism and Management Issues, chapter 15, pages 266 to 268, Cetaceans that are typically lonely and seek human company. Retrieved
December 17 2006.
Feeding
Various methods of feeding exist, not just between species but also within a species various methods may be employed, some techniques being used by only a single dolphin population. Fish and squid are the main source of food for most dolphin species, but the False Killer Whale and the Killer Whale also feed on other marine mammals.
One feeding method employed by many species is herding, where a pod will control a school of fish while individual members take turns plowing through the school, feeding. The tightly packed school of fish is commonly known as bait ball. Coralling is a method where fish are chased to shallow water where they are more easily captured. In
South Carolina, the Atlantic Bottlenose Dolphin takes this one step further with what has become known as strand feeding, where the fish are driven onto mud banks and retrieved from there. "Coastal Stock(s) of Atlantic Bottlenose Dolphin: Status Review and Management," Proceedings and Recommendations from a Workshop held in Beaufort, North Carolina, 13-14 September 1993. U.S. Department of Commerce, National Oceanic and Atmospheric Administration, National Marine Fisheries Service. pp. 56-57. In some places, Orcas will also come up to the beach to capture
sea lions. Some species also whack fish with their fluke, stunning them and sometimes sending fish clear out of the water.
Reports of cooperative human-dolphin fisheries date back to the ancient Rome author and natural philosophy
Pliny the Elder.M.B. Santos, R. Fernández, A. López, J.A. Martínez and G.J. Pierce (2007), Variability in the diet of bottlenose dolphin, Tursiops truncatus, in Galician waters, north-western Spain, 1990 – 2005 (.pdf), article retrieved April 3, 2007. A modern human-dolphin fishery still takes place in Laguna, Brazil, Santa Catarina (state),
Brazil.The Telegraph (2006), Brazil's sexiest secret, article retrieved March 11, 2007.
Vocalizations
Dolphins are capable of making a broad range of sounds using nasal airsacs located just below the blowhole. Roughly three categories of sounds can be identified however; frequency modulated sounds which are usually just called whistles; burst-pulsed sounds and clicks. Whistles are used by dolphins to communicate, though the nature and extent of their ability to communicate in this way is not known. Research has shown however that at least some dolphin species are capable of sending identity information to each other using a signature whistle; a whistle that refers specifically to the identity of a certain dolphin. The burst-pulsed sounds are also used for communication, but again the nature and extent of communication possible this way is not known. Atlantic Spotted Dolphin vocalizations, chapter Delphinid vocalizations., the dolphin communication project, article retrieved August 7, 2007. The clicks are directional and used by dolphins for echolocation and are often in a short series called a click train, the rate increasing when approaching an object of interest. Dolphin echolocation clicks are amongst the
Whale song#Toothed Whale Sound Levels in the sea.W. W. L. Au, The Sonar of Dolphins (Springer, NY, 1993).
Jumping and playing
Dolphins often leap above the water surface, sometimes performing acrobatic figures (e.g. the
spinner dolphin). Scientists are not always quite certain about the purpose of this behaviour and the reason for it may vary, it could be to locate schools of fish by looking at above-water signs like feeding birds, they could be communicating to other dolphins to join a hunt, attempting to dislodge parasites, or simply doing it for fun. Play is a very important part of dolphins' lives, and they can often be observed playing with seaweed or play-fighting with other dolphins. They even harass other locals, like seabirds and turtles. Dolphins also seem to enjoy riding waves and frequently 'surf' coastal swells and the bow waves of boats.
Threats to dolphins
Natural threats to dolphins
Except for mankind (discussed below), dolphins have few natural enemies, some species or specific populations having none at all making them apex predators. For most smaller species of dolphins, only a few larger species of shark such as the bull shark, dusky shark,
tiger shark and
great white shark are a potential risk, especially for calves. Some of the larger dolphin species such as Orcas may also prey on some of the smaller dolphin species, but this seems rare. Dolphins may also suffer from a wide variety of
diseases and parasites.
Human threats to dolphins
on the
Faroe Islands, killed in a drive hunt. Some dolphin species face an uncertain future, especially some of the river dolphin species such as the Amazon River dolphin, and the Ganges and Baiji, all of which are critically or seriously endangered. A 2006 survey found no individuals of the Yangtze River dolphin, leading to the conclusion that the species is now functionally extinct.Douglas Williams for Shanghai Daily (2006), Yangtze dolphin may be extinct. Article retrieved December 9
2006.
Contamination of environment - the oceans, seas, and rivers - is an issue of concern, especially
pesticides, heavy metals, plastics, and other industrial and agricultural pollutants which do not disintegrate rapidly in the environment are reducing dolphin populations, and resulting in dolphins building up unusually high levels of contaminants. Injuries or deaths due to collisions with boats, especially their
propellers, are also common.
Various fishing methods, most notably Seine (fishing) fishing for
tuna and the use of drift and gill nets, results in a large amounts of dolphins being killed inadvertently.Clover, Charles. 2004.
The End of the Line: How overfishing is changing the world and what we eat. Ebury Press, London. ISBN 0-09-189780-7 Accidental by-catch in trout nets is common and poses a risk for mainly local dolphin populations. In some parts of the world, such as some areas in Japan and the Faroe Islands, dolphins are traditionally considered as food, and killed in harpoon or
dolphin drive hunting.
Human-dolphin relationships
Mythology
(mount) Makara (Hindu mythology) Dolphins have long played a role in human culture. Dolphins are common in
Greek mythology and there are many coins from the time which feature a man or boy riding on the back of a dolphin. The Ancient Greece treated them with welcome; a ship spotting dolphins riding in their wake was considered a good omen for a smooth voyage. Dolphins also seem to have been important to the Minoans, judging by artistic evidence from the ruined palace at
Knossos. In
Hindu mythology, the Ganges River Dolphin is associated with Ganga, the deity of the
Ganges river.
Therapy
Dolphins are an increasingly popular choice of animal-assisted therapy for psychological problems and developmental disabilities. For example, a 2005 study with 30 participants found it was an effective treatment for mild to moderate
clinical depression.Christian Antonioli and Michael A. Reveley, (2005), Randomised controlled trial of animal facilitated therapy with dolphins in the treatment of depression. However, this study was criticized on several grounds; for example, it is not known whether dolphins are more effective than common pets.
Reviews of this and other published dolphin-assisted therapy (DAT) studies have found important methodological flaws and have concluded that there is no compelling scientific evidence that DAT is a legitimate therapy or that it affords any more than fleeting improvements in mood.
Entertainment
In more recent times, the Flipper (1963 film) and the subsequent popular
Flipper (1964 TV series), contributed to the popularity of dolphins in
Western World. The series, created by Ivan Tors, portrayed a dolphin in a friendly relationship with two boys, Sandy and Bud; a kind of seagoing
Lassie (1954 TV series). Flipper, a Bottlenose Dolphin, understood English unusually well and was a marked hero. A Flipper (1996 film) was made in
1996, which was based on the story of the original movie. A Bottlenose Dolphin also played a prominent role in the 1990s
science fiction television series
seaQuest DSV in which the animal, named Darwin, could communicate with English speakers using a vocoder, a fictional invention which translated the clicks and whistles to English and back., Dominican republic. More well known from this time period is probably the movie
Free Willy however, which made famous the Orca playing Willy, Keiko (orca). The 1977
horror movie Orca (film) paints a less friendly picture of the animal. Here, a male Orca takes revenge on fishermen after the killing of his mate. In the
1973 movie
The Day of the Dolphin trained dolphins are kidnapped and made to perform a naval military assassination using explosives. The renewed popularity of dolphins in the 1960s resulted in the appearance of many
dolphinariums around the world, which have made dolphins accessible to the public. Though criticism and more strict
animal welfare laws have forced many dolphinariums to close their doors, hundreds still exist around the world attracting a large amount of visitors. In the
United States, best known are the SeaWorld marine mammal parks, and their common Orca stage name
Shamu, which they have
trademarked, has become well known.
Southwest Airlines, an American airline, has even painted three of their Boeing 737 aircraft in
Shamu colours as an advertisement for the parks and have been flying with such a livery on various aircraft since
1988.
Occasionally, dolphins make an appearance in
computer games. Best known is the
Ecco the Dolphin game series. The games are named after their main character, Ecco, a young Bottlenose Dolphin. The Ecco the Dolphin games hinge on the idea that cetaceans are sapient beings and have their own underwater society.
A well known American National Football League (NFL) team is named the
Miami Dolphins. Their logo depicts an aqua-coloured Bottlenose Dolphin wearing an
American football helmet and jumping in front of a coral-coloured sunburst.
Military
A number of militaries have employed dolphins for various purposes from finding mines to rescuing lost or trapped humans. Such
military dolphins, however, drew scrutiny during the Vietnam War when rumors circulated that dolphins were being trained to kill Vietnamese Skin diving. Best known today is the U.S. Navy Marine Mammal Program.
Literature
Dolphins are also common in contemporary literature, especially science fiction novels. A military role for dolphins is found in William Gibson's short story
Johnny Mnemonic, in which cyborg dolphins are used in war-time by the military to find submarines and, after the war, by a group of revolutionaries to decode encrypted information. Dolphins play a role as sentient patrollers of the sea enhanced with a deeper empathy toward humans in
Anne McCaffrey's
Dragonriders of Pern series. In the
Known Space universe of author
Larry Niven, dolphins also play a significant role as fully-recognised "legal entities". More humorous is
The Hitchhiker's Guide to the Galaxy, in which dolphins are the second most intelligent creatures on Earth (after mice, and followed by humans) and tried in vain to warn humans of the impending destruction of the planet. However, their behaviour was misinterpreted as playful
acrobatics. Their story is told in
So Long, and Thanks for All the Fish. Much more serious is their major role (along with
chimpanzees) in David Brin's
Uplift Universe series. A talking Dolphin called "Howard" helps
Hagbard Celine and his submarine crew fight the evil
Illuminati in Robert Shea and Robert Anton Wilson's
Illuminatus Trilogy.
Dolphins also appear frequently in non-science fiction literature however. In the book
The Music of Dolphins by author Karen Hesse, a girl is raised by dolphins from the age of four until she is discovered by the
coast guard. Fantasy author
Ken Grimwood wrote dolphins into his
1995 novel
Into the Deep about a
marine biology struggling to crack the code of
cetacean intelligence, including entire chapters written from the viewpoint of his dolphin characters. In this book, humans and dolphins are capable of communicating via telepathy.
2007: Year of the Dolphin
The year
2007 has been declared as
(International) Year of the Dolphin by the United Nations and
United Nations Environment Programme (UNEP) in an effort to promote dolphin conservation. 2007: (International) Year of the Dolphin - Official website, website retrieved on January 14, 2007. The idea was launched by the UN's Convention on Migratory Species and the Whale and Dolphin Conservation Society (WDCS).
References
External links
Further information:
- OM Place - pictorial comparative chart of various dolphin species.
- Dolphins and their significance in world mythology.
- Tursi's dolphin page
Dolphin conservation and research:
- The Whale & Dolphin Conservation Society (WDCS)
- Charityguide.com - Save Bottlenose dolphins
- The Dolphin Institute
- The Dolphin research center
- Digital Library of Dolphin Development, Cetacean origins, Thewissen Lab
Dolphin news:
- Tursiops.org: Current Cetacean-related news
Dolphin photos:
- Red Sea Spinner Dolphin - Photo gallery
- PBS NOVA: Dolphins: Close Encounters
- David's Dolphin Images
- Images of Wild Dolphins in the Red Sea
- National Geographic
{{Taxobox| color = pink| name = Dolphin| image = Bottlenose_Dolphin_KSC04pd0178.jpg| image_width = 250px| image_caption = Bottlenose Dolphin breaching in the bow wave of a boat] - Recent| regnum = Animalia]| classis = Mammalia]| subordo = Odontoceti, 1821| subdivision_ranks = [Genus| subdivision =See article below.-->
Dolphins are
aquatic mammals that are closely related to whales and
porpoises. There are almost forty species of dolphin in seventeen
genus. They vary in size from 1.2 metres (4 foot (unit of length)) and 40 kilograms (88 pound (unit of weight)) (
Maui's Dolphin), up to 9.5 m (30 ft) and ten tonnes (the
Orca or Killer Whale). They are found worldwide, mostly in the shallower seas of the continental shelves, and are carnivores, mostly eating fish and squid. The Family (biology) Oceanic dolphin is the largest in the Cetacea, and relatively recent: dolphins evolved about ten million years ago, during the
Miocene. Dolphins are considered to be amongst the most intelligent of animals and their often friendly appearance and seemingly playful attitude have made them popular in human culture.
Origin of the name
The name is originally from
Ancient Greek (
delphís; "dolphin"), which was related to the Greek (
delphys; "womb"). The animal's name can therefore be interpreted as meaning "a 'fish' with a womb".The American Heritage® Dictionary of the English Language, Fourth Edition, online entry at Dictionary.com, retrieved December 17
2006. The name was transmitted via the Latin
delphinus, Middle Latin
dolfinus and the
Old French daulphin, which reintrodued the
ph into the word.
The word is used in a few different ways. It can mean:
- Any member of the family Delphinidae (oceanic dolphins),
- Any member of the families Delphinidae and Platanistoidea (oceanic and river dolphins),
- Any member of the suborder Odontoceti (toothed whales; these include the above families and some others),
- Used casually as a synonym for Bottlenose Dolphin, the most common and familiar species of dolphin.
In this article, the second definition is used.
Porpoises (suborder Odontoceti, family Phocoenidae) are thus not dolphins in this sense. Orcas and some closely related species belong to the Delphinidae family and therefore qualify as dolphins, even though they are called whales in common language. A group of dolphins can be called a "school" or a "pod".
Taxonomy
- Suborder Odontoceti, toothed whales
- Family Delphinidae, oceanic dolphins
- Genus Delphinus
- Genus Tursiops
- Genus Lissodelphis
- Northern Rightwhale Dolphin, Lissodelphis borealis
- Southern Rightwhale Dolphin, Lissiodelphis peronii
- Genus Sotalia
- Tucuxi, Sotalia fluviatilis
- Genus Sousa
- Genus Stenella
- Atlantic Spotted Dolphin, Stenella frontalis
- Clymene Dolphin, Stenella clymene
- Pantropical Spotted Dolphin, Stenella attenuata
- Spinner Dolphin, Stenella longirostris
- Striped Dolphin, Stenella coeruleoalba
- Genus Steno
- Rough-Toothed Dolphin, Steno bredanensis
- Genus Cephalorynchus
- Genus Grampus
- Risso's Dolphin, Grampus griseus
- Genus Lagenodelphis
- Genus Lagenorhyncus
- Genus Orcaella
- Genus Peponocephala
- Melon-headed Whale, Peponocephala electra
- Genus Orcinus
- Killer Whale, Orcinus orca
- Genus Feresa
- Pygmy Killer Whale, Feresa attenuata
- Genus Pseudorca
- Genus Globicephala
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhynchus
- Family Platanistoidea, river dolphins
- Genus Inia
- Boto (Amazon River Dolphin), Inia geoffrensis
- Genus Lipotes
- Genus Platanista
- Genus Pontoporia
Six species in the family Delphinidae are commonly called "whales" but are strictly speaking dolphins. They are sometimes called "blackfish".
- Melon-headed Whale, Peponocephala electra
- Orca, Orcinus orca
- Pygmy Killer Whale, Feresa attenuata
- False Killer Whale, Psudorca crassidens
- Long-finned Pilot Whale, Globicephala melas
- Short-finned Pilot Whale, Globicephala macrorhynchus
Hybrid dolphins
In 1933, three abnormal dolphins were beached off the Ireland coast; these appeared to be
hybrids between Risso's Dolphin and the Bottlenose Dolphin.Dolphin Safari (2006) sightings log, page retrieved December 17
2006. This mating has since been repeated in captivity and a hybrid calf was born. In captivity, a Bottlenose Dolphin and a Rough-toothed Dolphin produced hybrid offspring.Texas Tech University (1997), Mammals of Texas - Rough-toothed Dolphin, article retrieved December 8
2006. A Common-Bottlenose hybrid lives at SeaWorld California.Robin's Island Dolphins at SeaWorld California, page retrieved December 17
2006. Various other dolphin hybrids have also been reported in the wild, such as a Bottlenose-Atlantic Spotted hybrid.Denise L. Herzing, Kelly Moewe and Barbara J. Brunnick (2003), Interspecies interactions between Atlantic spotted dolphins, Stenella frontalis and bottlenose dolphins, Tursiops truncatus, on Great Bahama Bank, Bahamas, article retrieved on December 17 2006. The best known hybrid however is the Wolphin, a False Killer Whale-Bottlenose Dolphin hybrid. The Wolphin is a fertile hybrid, and two such Wolphins currently live at the Sea Life Park in Hawaii, the first having been born in 1985 from a male False Killer Whale and a female Bottlenose. Wolphins have also been observed in the wild.
Louis Herman, interviewed for Associated Press, article by Jeanette J. Lee (2005), Livescience.com - Whale-Dolphin Hybrid Has Baby Wholphin, article retrieved April 26, 2007.
Evolution and anatomy
Evolution
Dolphins, along with whales and porpoises, are thought to be descendants of terrestrial mammals, most likely of the
Artiodactyl order (biology). The ancestors of the modern day dolphins entered the water roughly fifty million years ago, in the
Eocene epoch.
Modern dolphin skeletons have two small, rod-shaped pelvic bones thought to be
vestigial hind legs. In
October 2006 an unusual Bottlenose Dolphin was captured in
Japan; it had small fins on each side of its genital slit which scientists believe to be a more pronounced development of these vestigial hind legs.
Associated Press /
FOX news (2006), Japanese Researchers Find Dolphin With 'Remains of Legs', article retrieved
November 6 2006.
Anatomy
Dolphins have a streamlined fusiform body, adapted for fast swimming. The basic colouration patterns are shades of grey with a light underside and a distinct dark cape on the back. It is often combined with lines and patches of different hue and contrast.
The head contains the
melon (whale), a round organ used for animal echolocation. In many species, the jaws are elongated, forming a distinct beak; for some species like the Bottlenose, there is a curved mouth which looks like a fixed smile. Teeth can be very numerous (up to two hundred and fifty) in several species. The dolphin brain is large and has a highly structured cortex, which often is referred to in discussions about their advanced intelligence.
Unlike most mammals, dolphins do not have hair, but they are born with a few hairs around the tip of their rostrum (anatomy) which they lose after some time, in some cases even before they are born. The only exception to this is the Boto river dolphin, which does have some small hairs on the rostrum.
Their reproductive organs are located on the underside of the body. Males have two slits, one concealing the penis and one further behind for the
anus. The female has one genital slit, housing the vagina and the anus. A mammary slit is positioned on either side of the female's genital slit.
Senses
Most dolphins have acute eyesight, both in and out of the water, and their sense of
Hearing (sense) is superior to that of humans. Though they have a small ear opening on each side of their head, it is believed that hearing underwater is also if not exclusively done with the lower jaw which conducts the sound vibrations to the middle ear via a fat-filled cavity in the lower jaw bone. Hearing is also used for Animal_echolocation, which seems to be an ability all dolphins have. Their teeth are arranged in a way that works as an array or antenna to receive the incoming sound and make it easier for them to pinpoint the exact location of an object.Goodson, A.D. and M. Klinowska. "A Proposed Echolocation Receptor for the Bottlenose Dolphin (
Tursiops truncatus): Modelling the Receive Directivity from Tooth and Lower Jaw Geometry", in Thomas and Kastelein, eds,
NATO ASI Series A: Sensory Abilities of Cetaceans, vi.196:255-267 (Plenum NY, 1990) ISBN 0-30-643695-7 The dolphin's sense of touch is also well-developed. However, dolphins lack an olfactory nerve and lobes and thus are believed to have no sense of smell,SeaWorld, Bottlenose Dolphins - Senses, article retrieved December 17 2006. but they can
taste and do show preferences for certain kinds of fish. Since dolphins spend most of their time below the surface normally, just tasting the water could act in a manner analogous to a sense of smell.
Though most dolphins do not have any hair, they do still have hair follicles and it is believed these might still perform some sensory function, though it is unclear what exactly this may be.Bjorn Mauck, Ulf Eysel and Guide Dehnhardt (2000), Selective heating of vibrissal follicles in seals (Phoca Vitulina) and dolphins (Sotalia Fluviatilis Guianensis), article retrieved March 11, 2007. The small hairs on the rostrum of the Boto river dolphin are believed to function as a
somatosensory system however, possibly to compensate for the Boto's poor eyesight.Laurie Stepanek (1998), Amazon River Dolphin (Inia geoffrensis), article retrieved
March 11, 2007.
Behaviour
Dolphins are often regarded as one of Earth's most intelligent animals, though it is hard to say just how intelligent dolphins are, as comparisons of species' relative intelligence are complicated by differences in sensory apparatus, response modes, and nature of cognition. Furthermore, the difficulty and expense of doing experimental work with large aquatics means that some tests which could yield meaningful results still have not been carried out, or have been carried out with inadequate sample size and methodology. Dolphin behaviour has been studied extensively by humans however, both in captivity and in the wild. See the
cetacean intelligence article for more details.
Social behaviour
, Queensland,
Australia.Dolphins are social, living in pods (also called "schools") of up to a dozen individuals. In places with a high abundance of food, pods can join temporarily, forming an aggregation called a
superpod; such groupings may exceed a thousand dolphins. The individuals communicate using a variety of clicks, whistles and other vocalizations. They also use ultrasonic sounds for echolocation. Membership in pods is not rigid; interchange is common. However, the cetaceans can establish strong bonds between each other. This leads to them staying with injured or ill individuals.
In May 2005, researchers in Australia discovered a cultural aspect of dolphin behaviour: Some dolphins (
Tursiops aduncus) teach their children to use tools. The dolphins break
sea sponges off and cover their snouts with them thus protecting their snouts while foraging. This knowledge of how to use a tool is mostly transferred from mothers to daughters, unlike simian
primates, where the knowledge is generally passed on to both sexes. The technology to use sponges as mouth protection is not genetically inherited but a taught behaviour.Rowan Hooper for New Scientist (2005), Dolphins teach their children to use sponges, article retrieved December 17
2006.
They are also occasionally willing to approach humans and playfully interact with them in the water. Dolphins have also been known to seemingly protect swimmers from sharks by swimming circles around them.CBC News (2004), Dolphins save swimmers from shark, article retrieved
March 11, 2007.
Dolphins are known to engage in acts of aggression towards each other. The older a male dolphin is, the more likely his body is covered with scars ranging in depth from teeth marks made by other dolphins. It is suggested that male dolphins engage in such acts of aggression for the same reasons as humans: disputes between companions or even competition for other females. Acts of aggression can become so intense that targeted dolphins are known to go into exile, leaving their communities as a result of losing a fight with other dolphins.
Male Bottlenose Dolphins have been known to engage in infanticide. Dolphins have also been known to kill porpoises for reasons which are not fully understood, as porpoises generally do not share the same fish diet as dolphins and are therefore not competitors for food supplies.Dr. George Johnson (date unknown), Is Flipper A Senseless Killer?, article retrieved December 17
2006.
Reproduction and sexuality
Dolphin copulation happens belly to belly and though many species engage in lengthy foreplay, the actual act is usually only brief, but may be repeated several times within a short timespan. The gestation period varies per species; for the small Tucuxi dolphin, this period is around 11 to 12 months, while for the Orca the gestation period is around 17 months. They usually become sexually active at a young age already, even before reaching sexual maturity. The age at which sexual maturity is reached varies per species and gender.
Dolphins are known to have sex for reasons other than reproduction, sometimes also engaging in acts of a homosexual nature.Herzing D.L., Rogers C.A., for the Wild Dolphin Project, Directionality of sexual aggression in mixed-species encounters between Atlantic Spotted dolphins and Bottlenose dolphins in the Bahamas (2005), article retrieved September 18, 2007. Various dolphin species have been known to engage in sexual behaviour with other dolphin species, this also having resulted in various hybrid dolphin species as mentioned earlier. Sexual encounters may be violent, with male dolphins sometimes showing aggressive behaviour towards both females and other male dolphins.Scott et. al, Aggression in bottlenose dolphins: evidence for sexual coercion, male-male competition, and female tolerance through analysis of tooth-rake marks and behaviour (2005), article retrieved September 18, 2007. Occasionally, dolphins will also show sexual behaviour towards other animals, including humans.Amy Samuels, Lars Bejder, Rochelle Constantine and Sonja Heinrich (2003), Marine Mammals: Fisheries, Tourism and Management Issues, chapter 15, pages 266 to 268, Cetaceans that are typically lonely and seek human company. Retrieved
December 17 2006.
Feeding
Various methods of feeding exist, not just between species but also within a species various methods may be employed, some techniques being used by only a single dolphin population. Fish and squid are the main source of food for most dolphin species, but the False Killer Whale and the Killer Whale also feed on other marine mammals.
One feeding method employed by many species is herding, where a pod will control a school of fish while individual members take turns plowing through the school, feeding. The tightly packed school of fish is commonly known as bait ball. Coralling is a method where fish are chased to shallow water where they are more easily captured. In
South Carolina, the Atlantic Bottlenose Dolphin takes this one step further with what has become known as strand feeding, where the fish are driven onto mud banks and retrieved from there. "Coastal Stock(s) of Atlantic Bottlenose Dolphin: Status Review and Management," Proceedings and Recommendations from a Workshop held in Beaufort, North Carolina, 13-14 September 1993. U.S. Department of Commerce, National Oceanic and Atmospheric Administration, National Marine Fisheries Service. pp. 56-57. In some places, Orcas will also come up to the beach to capture
sea lions. Some species also whack fish with their fluke, stunning them and sometimes sending fish clear out of the water.
Reports of cooperative human-dolphin fisheries date back to the
ancient Rome author and
natural philosophy Pliny the Elder.M.B. Santos, R. Fernández, A. López, J.A. Martínez and G.J. Pierce (2007), Variability in the diet of bottlenose dolphin, Tursiops truncatus, in Galician waters, north-western Spain, 1990 – 2005 (.pdf), article retrieved April 3, 2007. A modern human-dolphin fishery still takes place in
Laguna, Brazil,
Santa Catarina (state), Brazil.The Telegraph (2006), Brazil's sexiest secret, article retrieved
March 11, 2007.
Vocalizations
Dolphins are capable of making a broad range of sounds using nasal airsacs located just below the blowhole. Roughly three categories of sounds can be identified however;
frequency modulated sounds which are usually just called whistles; burst-pulsed sounds and clicks. Whistles are used by dolphins to communicate, though the nature and extent of their ability to communicate in this way is not known. Research has shown however that at least some dolphin species are capable of sending identity information to each other using a signature whistle; a whistle that refers specifically to the identity of a certain dolphin. The burst-pulsed sounds are also used for communication, but again the nature and extent of communication possible this way is not known. Atlantic Spotted Dolphin vocalizations, chapter Delphinid vocalizations., the dolphin communication project, article retrieved August 7, 2007. The clicks are directional and used by dolphins for echolocation and are often in a short series called a click train, the rate increasing when approaching an object of interest. Dolphin echolocation clicks are amongst the
Whale song#Toothed Whale Sound Levels in the sea.W. W. L. Au, The Sonar of Dolphins (Springer, NY, 1993).
Jumping and playing
Dolphins often leap above the water surface, sometimes performing acrobatic figures (e.g. the
spinner dolphin). Scientists are not always quite certain about the purpose of this behaviour and the reason for it may vary, it could be to locate schools of fish by looking at above-water signs like feeding birds, they could be communicating to other dolphins to join a hunt, attempting to dislodge parasites, or simply doing it for fun. Play is a very important part of dolphins' lives, and they can often be observed playing with seaweed or play-fighting with other dolphins. They even harass other locals, like seabirds and turtles. Dolphins also seem to enjoy riding waves and frequently 'surf' coastal swells and the bow waves of boats.
Threats to dolphins
Natural threats to dolphins
Except for mankind (discussed below), dolphins have few natural enemies, some species or specific populations having none at all making them
apex predators. For most smaller species of dolphins, only a few larger species of shark such as the
bull shark,
dusky shark, tiger shark and
great white shark are a potential risk, especially for calves. Some of the larger dolphin species such as Orcas may also prey on some of the smaller dolphin species, but this seems rare. Dolphins may also suffer from a wide variety of diseases and
parasites.
Human threats to dolphins
on the
Faroe Islands, killed in a drive hunt. Some dolphin species face an uncertain future, especially some of the river dolphin species such as the Amazon River dolphin, and the Ganges and
Baiji, all of which are critically or seriously endangered. A 2006 survey found no individuals of the Yangtze River dolphin, leading to the conclusion that the species is now
functionally extinct.Douglas Williams for Shanghai Daily (2006), Yangtze dolphin may be extinct. Article retrieved December 9 2006.
Contamination of environment - the oceans, seas, and rivers - is an issue of concern, especially
pesticides, heavy metals, plastics, and other industrial and agricultural pollutants which do not disintegrate rapidly in the environment are reducing dolphin populations, and resulting in dolphins building up unusually high levels of contaminants. Injuries or deaths due to collisions with boats, especially their propellers, are also common.
Various fishing methods, most notably Seine (fishing) fishing for
tuna and the use of drift and gill nets, results in a large amounts of dolphins being killed inadvertently.Clover, Charles. 2004.
The End of the Line: How overfishing is changing the world and what we eat. Ebury Press, London. ISBN 0-09-189780-7 Accidental by-catch in
trout nets is common and poses a risk for mainly local dolphin populations. In some parts of the world, such as some areas in Japan and the Faroe Islands, dolphins are traditionally considered as food, and killed in harpoon or
dolphin drive hunting.
Human-dolphin relationships
Mythology
(mount) Makara (Hindu mythology) Dolphins have long played a role in human culture. Dolphins are common in Greek mythology and there are many coins from the time which feature a man or boy riding on the back of a dolphin. The Ancient Greece treated them with welcome; a ship spotting dolphins riding in their wake was considered a good omen for a smooth voyage. Dolphins also seem to have been important to the Minoans, judging by artistic evidence from the ruined palace at
Knossos. In
Hindu mythology, the Ganges River Dolphin is associated with
Ganga, the deity of the
Ganges river.
Therapy
Dolphins are an increasingly popular choice of
animal-assisted therapy for psychological problems and developmental disabilities. For example, a 2005 study with 30 participants found it was an effective treatment for mild to moderate
clinical depression.Christian Antonioli and Michael A. Reveley, (2005), Randomised controlled trial of animal facilitated therapy with dolphins in the treatment of depression. However, this study was criticized on several grounds; for example, it is not known whether dolphins are more effective than common pets. Reviews of this and other published dolphin-assisted therapy (DAT) studies have found important methodological flaws and have concluded that there is no compelling scientific evidence that DAT is a legitimate therapy or that it affords any more than fleeting improvements in mood.
Entertainment
In more recent times, the Flipper (1963 film) and the subsequent popular
Flipper (1964 TV series), contributed to the popularity of dolphins in Western World. The series, created by
Ivan Tors, portrayed a dolphin in a friendly relationship with two boys, Sandy and Bud; a kind of seagoing
Lassie (1954 TV series). Flipper, a Bottlenose Dolphin, understood English unusually well and was a marked hero. A Flipper (1996 film) was made in 1996, which was based on the story of the original movie. A Bottlenose Dolphin also played a prominent role in the 1990s science fiction television series
seaQuest DSV in which the animal, named Darwin, could communicate with English speakers using a vocoder, a fictional invention which translated the clicks and whistles to English and back., Dominican republic. More well known from this time period is probably the movie
Free Willy however, which made famous the Orca playing Willy,
Keiko (orca). The 1977 horror movie
Orca (film) paints a less friendly picture of the animal. Here, a male Orca takes revenge on fishermen after the killing of his mate. In the
1973 movie
The Day of the Dolphin trained dolphins are kidnapped and made to perform a naval military assassination using explosives. The renewed popularity of dolphins in the 1960s resulted in the appearance of many dolphinariums around the world, which have made dolphins accessible to the public. Though criticism and more strict
animal welfare laws have forced many dolphinariums to close their doors, hundreds still exist around the world attracting a large amount of visitors. In the
United States, best known are the SeaWorld marine mammal parks, and their common Orca
stage name Shamu, which they have
trademarked, has become well known.
Southwest Airlines, an American airline, has even painted three of their
Boeing 737 aircraft in
Shamu colours as an advertisement for the parks and have been flying with such a livery on various aircraft since
1988.
Occasionally, dolphins make an appearance in
computer games. Best known is the
Ecco the Dolphin game series. The games are named after their main character, Ecco, a young Bottlenose Dolphin. The Ecco the Dolphin games hinge on the idea that cetaceans are sapient beings and have their own underwater society.
A well known American
National Football League (NFL) team is named the Miami Dolphins. Their logo depicts an aqua-coloured Bottlenose Dolphin wearing an
American football helmet and jumping in front of a coral-coloured sunburst.
Military
A number of militaries have employed dolphins for various purposes from finding mines to rescuing lost or trapped humans. Such military dolphins, however, drew scrutiny during the Vietnam War when rumors circulated that dolphins were being trained to kill Vietnamese
Skin diving. Best known today is the U.S. Navy Marine Mammal Program.
Literature
Dolphins are also common in contemporary literature, especially science fiction novels. A military role for dolphins is found in William Gibson's short story
Johnny Mnemonic, in which cyborg dolphins are used in war-time by the military to find submarines and, after the war, by a group of revolutionaries to decode encrypted information. Dolphins play a role as sentient patrollers of the sea enhanced with a deeper empathy toward humans in Anne McCaffrey's
Dragonriders of Pern series. In the
Known Space universe of author
Larry Niven, dolphins also play a significant role as fully-recognised "legal entities". More humorous is
The Hitchhiker's Guide to the Galaxy, in which dolphins are the second most intelligent creatures on Earth (after mice, and followed by humans) and tried in vain to warn humans of the impending destruction of the planet. However, their behaviour was misinterpreted as playful acrobatics. Their story is told in
So Long, and Thanks for All the Fish. Much more serious is their major role (along with chimpanzees) in
David Brin's
Uplift Universe series. A talking Dolphin called "Howard" helps
Hagbard Celine and his submarine crew fight the evil
Illuminati in Robert Shea and
Robert Anton Wilson's
Illuminatus Trilogy.
Dolphins also appear frequently in non-science fiction literature however. In the book
The Music of Dolphins by author
Karen Hesse, a girl is raised by dolphins from the age of four until she is discovered by the
coast guard. Fantasy author
Ken Grimwood wrote dolphins into his 1995 novel
Into the Deep about a marine biology struggling to crack the code of cetacean intelligence, including entire chapters written from the viewpoint of his dolphin characters. In this book, humans and dolphins are capable of communicating via telepathy.
2007: Year of the Dolphin
The year
2007 has been declared as
(International) Year of the Dolphin by the United Nations and United Nations Environment Programme (
UNEP) in an effort to promote dolphin conservation. 2007: (International) Year of the Dolphin - Official website, website retrieved on January 14, 2007. The idea was launched by the UN's
Convention on Migratory Species and the
Whale and Dolphin Conservation Society (WDCS).
References
External links
Further information:
- OM Place - pictorial comparative chart of various dolphin species.
- Dolphins and their significance in world mythology.
- Tursi's dolphin page
Dolphin conservation and research:
- The Whale & Dolphin Conservation Society (WDCS)
- Charityguide.com - Save Bottlenose dolphins
- The Dolphin Institute
- The Dolphin research center
- Digital Library of Dolphin Development, Cetacean origins, Thewissen Lab
Dolphin news:
- Tursiops.org: Current Cetacean-related news
Dolphin photos:
- Red Sea Spinner Dolphin - Photo gallery
- PBS NOVA: Dolphins: Close Encounters
- David's Dolphin Images
- Images of Wild Dolphins in the Red Sea
- National Geographic
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